1. Department of Mountain and Environmental Science, Interdisciplinary Graduate School of Science and Technology, Shinshu University, Asahi, Matsumoto, Nagano, Japan
2. Department of Biology, Faculty of Science, Shinshu University, Asahi, Matsumoto, Nagano, Japan
3. Institute of Mountain Science, Shinshu University, Asahi, Matsumoto, Nagano, Japan