1. Key Laboratory of Orogenic Belts and Crustal Evolution, School of Earth and Space Sciences, Peking University , 5 Yiheyuan Road, Beijing, 100871 , China
2. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences , Beijing, 10044 , China
3. CAS Center for Excellence in Life and Paleoenvironment , Beijing, 100044 , China
4. University of Chinese Academy of Sciences , Beijing, 100049 , China
5. Institut Català de Paleontologia, Universitat Autònoma de Barcelona , Cerdanyola del Vallès, 08193 Barcelona , Spain
6. Hezheng Paleozoological Museum , Hezheng, 731200 , China