1. Centre for Integrative Ecology, School of Life and Environmental Sciences, Deakin University , Waurn Ponds 3220 , Australia
2. Komodo Survival Program , Denpasar, Bali , Indonesia
3. Department of Biology, University of Florence , 50019 Sesto Fiorentino , Florence, Italy
4. Komodo National Park , Labuan Bajo, Flores , Indonesia