1. Biomedical Research Centre, Ninewells Hospital and Medical School, University of Dundee, Dundee DD1 9SY, Scotland, United Kingdom
2. Disease Mechanisms and Therapeutics Research Group, Department of Biological Sciences, University of Essex, Essex CO4 3SQ, United Kingdom
3. Centre for Tsukuba Advanced Research Alliance and Institute of Basic Medical Sciences, University of Tsukuba, Tsukuba 305-8577, Japan