1. SynthSys & Institute of Molecular Plant Sciences, School of Biological Sciences, King’s Buildings, University of Edinburgh , Edingburgh EH9 3BF , UK
2. School of Biological Sciences, University of Bristol , 24 Tyndall Avenue, Bristol BS8 1TQ , UK
3. Department of Cell and Molecular Biology, Uppsala University , S-751 24 Uppsala , Sweden
4. Plant Biology Section, School of Integrative Plant Science, Cornell University , Ithaca, NY , USA