1. Wellcome Centre for Cell Biology, University of Edinburgh , Michael Swann Building Max Born Crescent, King’s Buildings, Edinburgh, EH9 3BF , UK
2. Centre for Cardiovascular Science, The Queen’s Medical Research Institute (QMRI), University of Edinburgh , BioQuarter, 47 Little France Crescent, Edinburgh, EH16 4TJ , UK
3. Centre for Cardiovascular Science, The Queen’s Medical Research Institute (QMRI), University of Edinburgh , BioQuarter, 47 Little France Crescent , Edinburgh, EH16 4TJ, UK