1. Department of Physiology, Veterinary School, Complutense University of Madrid , Avenida Puerta de Hierro s/n, 28040 Madrid , Spain
2. Department of Integrative Biology and Evolution, Konrad Lorenz Institute of Ethology (KLIVV), University of Veterinary Medicine Vienna , Savoyenstraße 1a, A-1160 Vienna , Austria
3. Dipartimento di Biologia, Università di Padova , Via U. Bassi 58/B, I-35131 Padova , Italy