1. Ministry of Education Key Laboratory for Biodiversity and Ecological Engineering, College of Life Sciences, Beijing Normal University , Beijing , China
2. College of Forestry/Wuzhishan National Long Term Forest Ecosystem Monitoring Research Station, Hainan University , Haikou , China
3. Ministry of Education Key Laboratory for Ecology of Tropical Islands, College of Life Sciences, Hainan Normal University , Haikou , China