1. Key Laboratory of Freshwater Fish Reproduction and Development (Ministry of Education), School of Life Sciences, Southwest University , Chongqing 400715 , China
2. Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Southwest Forestry University , Kunming 650224 , China
3. School of Life Sciences, Yunnan University , Kunming 650091 , China
4. School of Ecology and Environment Sciences, Yunnan University , Kunming 650091 , China
5. State Key Laboratory of Genetic Resources and Evolution and Yunnan Key Laboratory of Biodiversity and Ecological Conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences , Kunming 650223 , China
6. Department of Ecology and Evolutionary Biology, University of Toronto , Toronto, Ontario , Canada