1. School of Life Sciences, Zhengzhou University , Zhengzhou, 450001 , China
2. Institute of Biodiversity and Ecology, Zhengzhou University , Zhengzhou, 450001 , China
3. School of Human Sciences, The University of Western Australia , WA 6009 , Australia
4. Centre for Evolutionary Biology, School of Biological Sciences, The University of Western Australia , WA 6009 , Australia
5. International Center of Biodiversity and Primate Conservation, Dali University , Dali, 671003 , China
6. Taihangshan Macaque Research Center , Jiyuan, 459000 , China