1. Institute of Transformative Bio-Molecules (WPI-ITbM), Nagoya University, Furo-cho, Chikusa-ku, Nagoya, Aichi 464-8601, Japan
2. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan
3. Center for Organismal Studies (COS), University of Heidelberg, Im Neuenheimer Feld 230, Heidelberg 69120, Germany
4. Cluster of Excellence on Plant Sciences, Heinrich Heine University, Universitätsstraße 1, Düsseldorf 40225, Germany