1. Machine Learning and Computational Biology Lab, Department of Biosystems Science and Engineering, ETH Zurich , 4058 Basel, Switzerland
2. Swiss Institute for Bioinformatics (SIB) , 1015 Lausanne, Switzerland
3. Google Research, Brain Team , 8002 Zürich, Switzerland
4. Institute of Mathematics and Statistics, University of Tartu , 51009 Tartu, Estonia
5. Institute of Genomics, University of Tartu , 51010 Tartu, Estonia
6. Department of Machine Learning and Systems Biology, Max Planck Institute of Biochemistry , 82152 Martinsried, Germany