1. Waseda Institute for Advanced Study, Waseda University , Tokyo 1690051, Japan
2. Department of Computer Science, Graduate School of Information Science and Technology, The University of Tokyo , Tokyo 1130032, Japan
3. Department of Integrated Biosciences, Graduate School of Frontier Sciences, The University of Tokyo , Chiba 2770882, Japan
4. Department of Biological Sciences, Graduate School of Science, The University of Tokyo , Tokyo 1130032, Japan
5. Department of Computational Biology and Medical Sciences, Graduate School of Frontier Sciences, The University of Tokyo , Chiba 2770882, Japan
6. Atmosphere and Ocean Research Institute, The University of Tokyo , Chiba 2770882, Japan
7. Institute for Quantitative Biosciences, The University of Tokyo , Tokyo 1130032, Japan
8. Collaborative Research Institute for Innovative Microbiology, The University of Tokyo , Tokyo 1130032, Japan