1. McDonnell Genome Institute, Washington University School of Medicine, St. Louis, MO 63108, USA
2. Department of Genetics, Washington University School of Medicine, St. Louis, MO 63110, USA
3. Biofrontiers Institute, University of Colorado, Boulder, CO 80309, USA
4. Department of Computer Science, University of Colorado, Boulder, CO 80309, USA
5. Department of Medicine, Washington University School of Medicine, St. Louis, MO 63110, USA