1. San Diego , CA 92172-721234, United States
2. Departments of Medicine and Bioengineering, University of Pittsburgh , Pittsburgh, PA 15213, United States
3. University of Pittsburgh Hillman Cancer Center , Pittsburgh, PA 15213, United States
4. Department of Computational and Systems Biology, University of Pittsburgh , Pittsburgh, PA 15213, United States
5. Department of Physics and Astronomy, University of Pittsburgh , Pittsburgh, PA 15213, United States