1. Computational Biology Department, School of Computer Science, Carnegie Mellon University, Pittsburgh, PA, USA
2. School of Cellular and Molecular Medicine, University of Bristol, Bristol BS, UK
3. Departments of Biological Sciences, Biomedical Engineering, and Machine Learning, Carnegie Mellon University, Pittsburgh, PA, USA
4. Freiburg Institute for Advanced Studies and Faculty of Biology, Albert Ludwig University of Freiburg, Freiburg im Breisgau, Baden-Württemberg, Germany