1. Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Aberystwyth SY23 3PD, UK
2. Department of Computer Science, Aberystwyth University, Aberystwyth SY23 3DB, UK
3. Department of Mathematics, Aberystwyth University, Aberystwyth SY23 3BZ, UK
4. School of Biological Sciences, Queen’s University Belfast, Belfast BT7 1NN, UK