1. Department of Biosciences, Durham University, Stockton Road, Durham DH1 3LE, UK
2. New England Biolabs, 240 County Road, Ipswich, MA 01938, USA
3. Department of Applied Sciences, University of Northumbria, Newcastle Upon Tyne NE1 8ST, UK
4. Institute of Infection, Veterinary and Ecological Sciences, University of Liverpool, Liverpool L69 7ZB, UK