1. Naturalis Biodiversity Center, PO Box 9517, 2300 RA Leiden, The Netherlands
2. Grand Valley State University, 1 Campus Drive, Allendale, MI 49401, USA
3. Institute of Biology Leiden, Leiden University, Sylviusweg 72, 2333 CC Leiden, The Netherlands
4. University of Applied Sciences Leiden, Zernikedreef 11, 2300 AJ Leiden, The Netherlands
5. Uppsala University, Norbyvägen 18D, 75236 Uppsala, Sweden
6. The Natural History Museum, University of Oslo, PO Box 1172, 0318 Oslo, Norway
7. Swedish Museum of Natural History, Frescativägen 40, 114 18 Stockholm, Sweden
8. University of Vienna, Universitätsring 1, 1010 Wien, Austria,
9. Sabah Park Herbarium, PO Box 6, Kinabalu Park, Kundasang, Ranau, Sabah, Malaysia
10. Hortus Botanicus of Leiden University, Rapenburg 73, 2311 GJ Leiden, The Netherlands