1. School of Agriculture, Food and Ecosystem Sciences, Faculty of Science, The University of Melbourne , Parkville, Melbourne, VIC 3010 , Australia
2. School of Biology, Faculty of Biological Sciences, The University of Leeds , Leeds LS2 9JT , UK
3. Centre for Innovation and Excellence in Livestock , Heslington, York YO10 5DG , UK
4. ICAR-Central Institute of Freshwater Aquaculture , Bhubaneswar , India