1. College of Veterinary Medicine, Yangzhou University , Yangzhou 225009 , PR China
2. Jiangsu Co-innovation Center for Prevention and Control of Important Animal Infectious Diseases and Zoonoses , Yangzhou 225009 , PR China
3. Joint International Research Laboratory of Agriculture and Agri-Product Safety, The Ministry of Education of China, Yangzhou University , Yangzhou 225009, Jiangsu , PR China
4. College of Veterinary Medicine, Albutana University , Rufaa 22217, Al Jazirah , Sudan