1. Guizhou Academy of Forestry , Guiyang, Guizhou 550005 , China
2. School of Life Sciences, Guizhou Normal University , Guiyang, Guizhou 550025 , China
3. Xiangzhong Normal College for Preschool Education , Shaoyang, Hunan 422099 , China
4. College of Forestry, Guizhou University , Guiyang, Guizhou 550025 , China