1. Chongqing Key Laboratory for Innovative Application of Genetic Technology, College of Resources and Environment, Southwest University , Chongqing 400715 , China
2. Key Laboratory of Healthy Freshwater Aquaculture, Ministry of Agriculture and Rural Affairs; Key Laboratory of Fish Health and Nutrition of Zhejiang Province, Zhejiang Institute of Freshwater Fisheries , Huzhou 313001 , China
3. National Base of International S&T Collaboration on Water Environmental Monitoring and Simulation in TGR Region (WEMST, www.wemst.com), College of Resources and Environment, Southwest University , Chongqing 400716 , China