1. Key Laboratory of Biofuels, Qingdao Carbon One Biorefining Research Center, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences , Qingdao 266100 , China
2. Shandong Energy Institute , Qingdao 266100 , China
3. Qingdao New Energy Shandong Laboratory , Qingdao 266100 , China
4. University of Chinese Academy of Sciences , College of Life Sciences, Beijing, 100039 , China