1. The United Graduate School of Agricultural Sciences, Tottori University , Tottori 680-8553 , Japan
2. Faculty of Agriculture, Tottori University , 4-101, Koyama-cho Minami, Tottori 680-8553 , Japan
3. Department of Biological Sciences, College of Science, University of Santo Tomas , España Blvd. 1015, Manila , Philippines
4. International Platform for Dryland Research and Education, Tottori University , 1390 Hamasaka, Tottori 680-0001 , Japan