1. Laboratoire d'Ecologie, Systématique et Evolution, CNRS UMR, Université Paris-Sud, Batiment 362, Orsay Cedex, France
2. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, College of Life Sciences, Beijing Normal University, Beijing, China
3. Ministry of Education Key Laboratory for Ecology of Tropical Islands, College of Life Sciences, Hainan Normal University, Haikou, China
4. Department of Ecology, Federal University of Goias, Goiania, Brazil