1. Nanjing Agricultural University Jiangsu Engineering Research Center for Pear, State Key Laboratory of Crop Genetics and Germplasm Enhancement, , Nanjing 210095, China
2. Nanjing Agricultural University State Key Laboratory of Crop Genetics and Germplasm Enhancement, , Nanjing 210095, China
3. Liaoning Academy of Agricultural Sciences Insitute of Pomology, , Yingkou 115009, China