1. Department of Epidemiology and Population Health, Institute of Infection and Global Health, University of Liverpool, Liverpool CH64 7TE, UK
2. Institute of Veterinary Science, University of Liverpool, Liverpool CH64 7TE, UK
3. Department of Life Sciences, University of Roehampton, London SW15 4JD, UK
4. Department of Health Services Research, Institute of Psychology, Health and Society, University of Liverpool, Liverpool L69 7ZX, UK