1. Laboratory of Vaccine Materials and Laboratory of Gut Environmental System, Microbial Research Center for Health and Medicine, National Institutes of Biomedical Innovation, Health and Nutrition (NIBIOHN) , Osaka 567-0085 , Japan
2. Graduate School of Pharmaceutical Sciences, Osaka University , Suita, Osaka 565-0871 , Japan
3. Department of Chemistry, Graduate School of Science, Osaka University , Osaka 560-0043 , Japan
4. Collaborative Research Between NIBIOHN and Graduate School of Science, Forefront Research Center, Osaka University , Osaka 560-0043 , Japan
5. Graduate School of Medicine, Osaka University , Osaka 565-0871 , Japan
6. Department of Life Sciences, Laboratory of Functional Anatomy, School of Agriculture, Meiji University , Kanagawa 214-8571 , Japan
7. Division of Gastroenterology, Department of Medicine, University of California San Diego (UCSD) , San Diego, CA 92093 , USA
8. Chiba University (CU)-UCSD Center for Mucosal Immunology, Allergy and Vaccines (cMAV), UCSD , San Diego, CA 92093-0063 , USA
9. Future Medicine Education and Research Organization, Chiba University , Chiba 260-8670 , Japan
10. Department of Human Mucosal Vaccinology, Chiba University Hospital , Chiba 260-8677 , Japan
11. International Vaccine Design Center, The Institute of Medical Science, The University of Tokyo , Tokyo 108-8639 , Japan
12. Department of Microbiology and Immunology, Kobe University Graduate School of Medicine , Kobe 650-0017 , Japan
13. Research Organization for Nano and Life Innovation, Waseda University , Tokyo 162-0041 , Japan
14. Graduate School of Dentistry, Osaka University , Suita 565-0871 , Japan