1. Division of Innate Immunity, Department of Microbiology and Immunology, The Institute of Medical Science, The University of Tokyo , Tokyo , Japan
2. Division of Clinical Genome Research, The Institute of Medical Science, The University of Tokyo , Tokyo , Japan
3. Department of Dermatology, Kyoto University Graduate School of Medicine , Kyoto , Japan
4. Alliance Laboratory for Advanced Medical Research, Kyoto University Graduate School of Medicine , Kyoto , Japan
5. Laboratory of Immune Regulation, School of Life Sciences, Tokyo University of Pharmacy and Life Sciences , Tokyo 192-0392 , Japan
6. Skin Research Institute of Singapore (SRIS) and A*STAR Skin Research Labs (A*SRL), Agency for Science, Technology, and Research (A*STAR) , 8A Biomedical Grove, IMMUNOS Building, Biopolis , Singapore