1. Laboratory of Lymphocyte Differentiation, WPI Immunology Frontier Research Center, Osaka University , Osaka 565-0871 , Japan
2. Division of Microbiology and Immunology, Center for Infectious Disease Education and Research, Osaka University , Osaka 565-0871 , Japan
3. Laboratory of Lymphocyte Differentiation, RIKEN Center for Integrative Medical Sciences (IMS) , Yokohama, Kanagawa 230-0045 , Japan