1. Division of Immunobiology, Research Institute for Biomedical Sciences, Tokyo University of Science, Yamazaki 2669, Noda, Chiba 278-0022, Japan
2. Department of Pathology, Teikyo University Chiba Medical Center, Ichihara, Chiba 299-0111, Japan
3. Division of Mucosal Immunology, Department of Microbiology and Immunology and
4. International Research and Development Center for Mucosal Vaccines, The Institute of Medical Science, The University of Tokyo, Tokyo 108-8639, Japan
5. Department of Innovative Medicine, Graduate School of Medicine,
6. Institute for Global Prominent Research and
7. Departments of Mucosal Immunology and Immunology, Graduate School of Medicine, Chiba University, Chiba 260-8670, Japan