1. Ann Romney Center for Neurologic Diseases, Department of Neurology, Brigham and Women’s Hospital and Harvard Medical School, Boston, MA 02115, USA
2. Department of Neurology, Columbia University Medical Center, New York, NY 10032, USA
3. Department of Cell Biology, Harvard Medical School, Boston, MA 02115, USA
4. Whitehead Institute for Biomedical Research, Cambridge, MA 02142, USA
5. HHMI, Department of Biology, MIT, Cambridge, MA 02139, USA