1. Eunice Kennedy Shriver National Institute of Child Health and Human Development Division of Developmental Biology, , Bethesda, MD, 20892, USA
2. University of Pittsburgh Pittsburgh Institute for Neurodegenerative Diseases, , Pittsburgh, PA, 15260, USA
3. University of Pittsburgh Department of Neurology, , Pittsburgh, PA,15260, USA
4. Geriatric Research, Education, and Clinical Center, Pittsburgh VA Healthcare System , Pittsburgh, PA, 15240, USA