1. Centre for Organismal Studies, Heidelberg University, Heidelberg, Germany
2. Computational Molecular Evolution Group, Heidelberg Institute for Theoretical Studies, Heidelberg, Germany
3. Institute for Theoretical Informatics, Karlsruhe Institute of Technology, Karlsruhe, Germany
4. Institute of Biological and Chemical Systems, Karlsruhe Institute of Technology, Eggenstein-Leopoldshafen, Germany