1. Computational and Molecular Evolutionary Biology Research Group, School of Life Sciences, Faculty of Medicine and Health Sciences, University of Nottingham , Nottingham NG7 2RD , United Kingdom
2. Computational and Molecular Evolutionary Biology Research Group, School of Biology, Faculty of Biological Sciences, University of Leeds , Leeds LS2 9JT , United Kingdom
3. Institute for Global Food Security, School of Biological Sciences, Queen's University Belfast , Belfast BT9 5DL , United Kingdom