1. Department of Ecology, Tibetan Centre for Ecology and Conservation at Wuhan University—Tibet University, Hubei Key Laboratory of Cell Homeostasis, College of Life Sciences, Wuhan University, Wuhan, China
2. Institute of Evolution, University of Haifa, Haifa, Israel
3. State Key Laboratory of Grassland Agro-Ecosystem, Institute of Innovation Ecology, Lanzhou University, Lanzhou, China
4. Department of Zoology, Faculty of Science, University of South Bohemia, České Budějovice, Czech Republic
5. Monell Chemical Senses Center, Philadelphia, PA, USA
6. Research Center for Ecology, College of Science, Tibet University, Lhasa, China