1. Department of Biology, University of Fribourg , Chemin du Musée 10, 1700 Fribourg , Switzerland
2. Institute of Plant Sciences, University of Bern , Altenbergrain 21, 3013 Bern , Switzerland
3. Central European Institute of Technology, Masaryk University , 625 00 Brno , Czech Republic
4. Royal Botanic Gardens , Kew, Surrey TW9 3AB , UK