1. Department of Ecology and Evolution, University of Lausanne, Lausanne, Switzerland
2. Swiss Institute of Bioinformatics, Lausanne, Switzerland
3. Institute of Evolutionary Sciences, University of Montpellier, CNRS, IRD, EPHE, Montpellier, France
4. Johann-Friedrich-Blumenbach Institute of Zoology and Anthropology, University of Goettingen, Goettingen, Germany