1. Department of Ecology and Evolutionary Biology, University of Toronto , Toronto, ON , Canada
2. Department of Cell and Systems Biology, University of Toronto , Toronto, ON , Canada
3. Department of Biological Sciences, Vanderbilt University , Nashville, TN
4. Department of Ophthalmology and Visual Sciences, Vanderbilt University Medical Center , Nashville, TN
5. Evolutionary Studies, Vanderbilt University , Nashville, TN
6. Department of Biological Sciences, University of Toronto Scarborough , Toronto, ON , Canada