1. School of Life Sciences, Arizona State University , Tempe, AZ 85287 , USA
2. Department of Biological Sciences, Mississippi State University , Mississippi State, MS 39762 , USA
3. Department of Biology, Indiana University , Bloomington, IN 47405 , USA
4. National Center for Genome Analysis Support, Indiana University , Bloomington, IN 47405 , USA
5. Center for Mechanisms of Evolution, The Biodesign Institute, Arizona State University , Tempe, AZ 85287 , USA