1. Graduate School of Human and Environmental Studies, Kyoto University, Yoshida nihonmatsu cho, Sakyo ku, Kyoto, Japan
2. Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic
3. Department of Biological Sciences, Mississippi State University, Mississippi State, MS, USA
4. Graduate School of Science, Kobe University, Hyogo, Japan
5. Japan Agency for Marine-Earth Science and Technology, Yokosuka, Japan
6. Graduate School of Agriculture, Kyoto University, Kitashirakawa oiwake cho, Sakyo ku, Kyoto, Japan