1. Laboratory for Neural Computation and Adaptation, RIKEN Center for Brain Science , 2-1 Hirosawa, Wako, Saitama 351-0106 , Japan
2. Theoretical Molecular Science Laboratory, RIKEN Cluster for Pioneering Research , 2-1 Hirosawa, Wako, Saitama 351-0198 , Japan
3. School of Physics, National Laboratory of Solid State Microstructure, and Collaborative Innovation Center of Advanced Microstructures, Nanjing University , Nanjing 210093 , People’s Republic of China
4. Center for Complex Systems Biology, Universal Biology Institute, University of Tokyo , Komaba, Meguro, Tokyo 153-8902 , Japan
5. The Niels Bohr Institute, University of Copenhagen , Blegdamsvej 17, Copenhagen 2100-DK , Denmark