1. Department of Environmental Sciences, Zoology, University of Basel , Basel 4051 , Switzerland
2. CEFE, Université de Montpellier, CNRS, EPHE, IRD , Montpellier 34293 , France
3. Tvärminne Zoological Station, University of Helsinki , Hanko 10900 , Finland
4. School of Zoology, George S. Wise Faculty of Life Sciences, Tel Aviv University , Tel Aviv 6997801 , Israel