1. Key Laboratory of Molecular Epigenetics of the Ministry of Education (MOE), Northeast Normal University , Changchun, Jilin 130024 , China
2. Jilin Academy of Vegetable and Flower Science , Changchun, Jilin 130033 , China
3. Laboratory of Plant Epigenetics and Evolution, School of Life Science, Liaoning University , Shenyang, Liaoning 110036 , China
4. Department of Ecology, Evolution and Organismal Biology, Iowa State University , Ames, IA 50010 , USA