1. Department of Research and Drug Discovery, Medical Innovation Center, Kyoto University Graduate School of Medicine, Kyoto, Japan
2. Department of Psychiatry and Behavioral Sciences, Johns Hopkins University School of Medicine, Baltimore, MD, USA
3. Faculty of Medicine, University of Tsukuba, Tsukuba 305-8575, Japan
4. CREST Project, Medical Innovation Center, Kyoto University Graduate School of Medicine, Kyoto, Japan
5. Department of Pharmacology, Kobe University Graduate School of Medicine, Hyogo, Japan
6. Department of Biophysics, Kyoto University Graduate School of Science, Kyoto, Japan
7. Campbell Family Mental Health Research Institute, Centre for Addiction and Mental Health, University of Toronto, Toronto, ON, Canada