1. Wellcome/CRC Institute Tennis Court Road Cambridge CB2 1QR UK
2. Department of Zoology, University of Cambridge Cambridge CB2 3EJ UK
3. Division of Molecular Cell Pharmacology, National Children's Medical Research Center 3‐35‐31 Taishido Setagayaku Tokyo 154‐8509 Japan
4. Department of Pathology, University of Cambridge Cambridge CB2 1QP UK