1. Key Laboratory of Plant Resource Conservation and Sustainable Utilization, South China Botanical Garden, The Chinese Academy of Sciences , Guangzhou 510650, China
2. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, The Chinese Academy of Sciences , Guangzhou 510650, China
3. School of Ecological and Environmental Sciences, Tiantong National Station for Forest Ecosystem Research, East China Normal University , Shanghai 200241, China