1. Microbiology, School of Natural Sciences, National University of Ireland, Galway, H91 TK33, Ireland
2. Information and Computational Sciences, James Hutton Institute, Invergowrie, Dundee DD2 5DA, Scotland
3. Soil and Environmental Microbiology, Environmental Research Centre, Teagasc, Johnstown Castle, Wexford, Y35 TC97, Ireland
4. Cell and Molecular Sciences, James Hutton Institute, Invergowrie, Dundee DD2 5DA, Scotland