1. Department of Life Sciences , Natural History Museum, Cromwell Road, London SW7 5BD , UK
2. Herforder Str. 190, Leopoldshöhe , North Rhine-Westphalia 33818 , Germany
3. Institute of Evolutionary Biology, Faculty of Biology, Biological and Chemical Research Centre, University of Warsaw , Żwirki i Wigury 101, Warsaw 02-089 , Poland